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http://onlinelibrary.wiley.com/doi/10.1002/1097-0185(20000701)259:3%3C312::AID-AR80%3E3.0.CO;2-X/full
Anchor text: Histo-physiology of the scent-marking glands of the penile pad, anal pouch, and the forefoot in the aardwolf (Proteles cristatus)